Showing posts with label DINOSAUR. Show all posts
Showing posts with label DINOSAUR. Show all posts

Monday 5 February 2024

CARNOTAURUS SASTREI: FLESH EATING BULL

Carnotaurus sastrei, a genus of large theropod dinosaurs that roamed the southern tip of Argentina, South America during the Late Cretaceous, 72 to 69.9 million years ago. His name means "flesh-eating bull,' and he lives up to it.

This fellow — or at least his robust skull with the short, knobby eyebrow horns and fierce-looking teeth — is on display at the Natural History Museum in Madrid, Spain. For now, he is the only known genus of this species of bipedal predator.

The first specimen of Carnotaurus sastrei was found in Chubut on vast plains between the Andes Mountains and the Atlantic Ocean. A physician, Dr. A'ngel Tailor noticed a large concretion showing some bone fragments. A team led by José F. Bonaparte excavated the find in 1984 as part of a paleontological expedition funded by the Argentine Museum of Natural Sciences.

Sadly, Bonaparte — the Maestro del Mesozoico — passed away the 18th February 20220 at the age of 91. He spent the majority of his career as head of the Vertebrate Palaeontology Division of the Museo Argentino de Ciencias Naturales “Bernardino Rivadavia,” in Buenos Aires. Bonaparte opened up the vertebrate finds of Argentina to the world. He was instrumental in the finding, excavating and naming many iconic dinosaurs — Carnotaurus, Amargasaurus, Abelisaurus, Argentinosaurus, Noasaurus along with the finding of the first fossilised remains of Mesozoic South American mammals. He mentored many palaeontologists who will miss his keen eye and tremendous work ethic — Luis Chiappe, Rodolfo Coria, Agustín Martinelli, Fernando Novas, Jaime Powell, Guillermo Rougier, Leonardo Salgado, Sebastián Apesteguía and many others.

His excavation of Carnotaurus was the first of its kind and he recognized that the skull is quite unusual. Initially, it has a very marine reptile feel — but make no mistake this guy is clearly a terrestrial theropod. He had smallish, underdeveloped arms — teeny by theropod standards. Once you look closer you see his bull-like horns from whence he gets his name — horns that imply battle between rivals for the best meal, sexual partner and to be the one who leads the herd. 

He was covered in leathery skin lined with rows of cone-shaped nodules or bumps. These get larger as they move towards his spine. He had forward-facing eyes, similar to tyrannosaurs like T-rex and smaller theropods like Velociraptor and Troodon — who had better vision even that T-rex — which would have given him the advantage of binocular vision and depth perception. Forward-facing eyes are also quite helpful with nocturnal hunting — think owls and cats — as they take in more light and help with nighttime predation. So perhaps this flesh-eating bull fancied a late-night snack on his menu from time to time.

Species like squirrels, pigeons and crocodiles have eyes on the sides of their heads. They lack the important competitive feature of well-developed depth perception — being able to easily and estimate distance — but perhaps make up for it with a panorama that offers a wider field of view.   

Friday 26 January 2024

FIRST DINOSAUR FROM VANCOUVER ISLAND

This dapper fellow is a pine needle and horsetail connoisseur. He's a hadrosaurus — a duck-billed dinosaur. They were a very successful group of plant-eaters that thrived throughout western Canada during the late Cretaceous, some 70 to 84 million years ago.

Hadrosaurs lived as part of a herd, dining on pine needles, horsetails, twigs and flowering plants.

Hadrosaurs are ornithischians — an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. They are close relatives and possibly descendants of the earlier iguanodontid dinosaurs. 

They had slightly webbed, camel-like feet with pads on the bottom for cushioning and perhaps a bit of extra propulsion in water. They were primarily terrestrial but did enjoy feeding on plants near and in shallow water. There had a sturdy build with a stiff tail and robust bone structure. 

At their emergence in the fossil record, they were quite small, roughly three meters long. That's slightly smaller than an American bison. They evolved during the Cretaceous with some of their lineage reaching up to 20 meters or 65 feet.

Hadrosaurs are very rare in British Columbia but a common fossil in our provincial neighbour, Alberta, to the east. Here, along with the rest of the world, they were more abundant than sauropods and a relatively common fossil find. They were common in the Upper Cretaceous of Europe, Asia, and North America.

There are two main groups of Hadrosaurs, crested and non-crested. The bony crest on the top of the head of the hadrosaurs was hollow and attached to the nasal passages. It is thought that the hollow crest was used to make different sounds. These sounds may have signalled distress or been the hadrosaur equivalent of a wolf whistle used to attract mates. Given their size it would have made for quite the trumpeting sound.

This beautiful specimen graces the back galleries of the Courtenay and District Museum on Vancouver Island, British Columbia, Canada. I was very fortunate to have a tour this past summer with the deeply awesome Mike Trask joined by the lovely Lori Vesper. The museum houses an extensive collection of palaeontological and archaeological material found on Vancouver Island, many of which have been donated by the Vancouver Island Palaeontological Society.

Dan Bowen, Chair of the Vancouver Island Palaeontological Society, shared the photo you see here of the first partly articulated dinosaur from Vancouver Island ever found. The vertebrate photo and illustration are from a presentation by Dr. David Evans at the 2018 Paleontological Symposium in Courtenay.  The research efforts of the VIPS run deep in British Columbia and this new very significant find is no exception. A Hadrosauroid dinosaur is a rare occurrence and further evidence of the terrestrial influence in the Upper Cretaceous, Nanaimo Group, Vancouver Island — outcrops that we traditionally thought of as marine from years of collecting well-preserved marine fossil fauna.

CDM 002 / Hadrosauroid Caudal Vertebrae
The fossil bone material was found years ago by Mike Trask of the Vancouver Island Palaeontological Society. You may recall that he was the same fellow who found the Courtenay Elasmosaur on the Puntledge River.

Mike was leading a fossil expedition on the Trent River. While searching through the Upper Cretaceous shales, the group found an articulated mass of bones that looked quite promising.

Given the history of the finds in the area, the bones were thought to be from a marine reptile.

Since that time, we've found a wonderful terrestrial helochelydrid turtle, Naomichelys speciosa, but up to this point, the Trent had been known for its fossil marine fauna, not terrestrial. Efforts were made to excavate more of the specimen, and in all more than 25 associated vertebrae were collected with the help of some 40+ volunteers. Identifying fossil bone is a tricky business. Encased in rock, the caudal vertebrae were thought to be marine reptile in origin. Some of these were put on display in the Courtenay Museum and mislabeled for years as an unidentified plesiosaur.

In 2016, after years of collecting dust and praise in equal measure, the bones were reexamined. They didn't quite match what we'd expect from a marine reptile. Shino Sugimoto, Fossil Preparator, Vertebrate Palaeontology Technician at the Royal Ontario Museum was called in to work her magic — painstakingly prepping out each caudal vertebrae from the block.

Once fully prepped, seemingly unlikely, they turned out to be from a terrestrial hadrosauroid. This is the second confirmed dinosaur from the Upper Cretaceous Nanaimo Group. The first being a theropod from Sucia Island consisting of a partial left thigh bone — the first dinosaur fossil ever found in Washington state.

Dr. David Evans, Temerty Chair in Vertebrate Palaeontology, Department of Natural History, Palaeobiology from the Royal Ontario Museum, confirmed the ID and began working on the partial duck-billed dinosaur skeleton to publish on the find.

Drawing of Trent River Hadrosauroid Caudal Vertebrae
Now fully prepped, the details of this articulated Hadrosauriod caudal vertebrae come to light. We can see the prominent chevron facets indicative of caudal vertebrae with a nice hexagonal centrum shape on its anterior view.

There are well-defined long, raked neural spines that expand distally — up and away from the acoelous centrum. 

Between the successive vertebrae, there would likely have been a fibrocartilaginous intervertebral body with a gel-like core —  the nucleus pulposus — which is derived from the embryonic notochord. This is a handy feature in a vertebrate built as sturdily as a hadrosaur. Acoelous vertebrae have evolved to be especially well-suited to receive and distribute compressive forces within the vertebral column.

This fellow has kissing cousins over in the state of New Jersey where this species is the official state fossil. The first of his kind was found by John Estaugh Hopkins in New Jersey back in 1838. Since that time, we've found many hadrosaurs in Alberta, particularly the Edmontosuaurs, another member of the subfamily Hadrosaurine.

In 1978, Princeton University found fifteen juvenile hadrosaurs, Maiasaura ("good mother lizard") on a paleontological expedition to the Upper Cretaceous, Two Medicine Formation of Teton County in western Montana. 

Their initial finds of several small skeletons had them on the hunt for potential nests — and they found them complete with wee baby hatchlings!

Photo One: Fossil Huntress / Heidi Henderson, VIPS

Photo Two / Sketch Three: Danielle Dufault, Palaeo-Scientific Ilustrator, Research Assistant at the Royal Ontario Museum, Host of Animalogic. 

The vertebrate photo and illustration were included in a presentation by Dr. David Evans at the 2018 BCPA Paleontological Symposium in Courtenay, British Columbia, Canada.

Photo Four: Illustration by the talented Greer Stothers, Illustrator & Natural Science-Enthusiast.

Wednesday 17 January 2024

COAHUILACERATOPS MAGNACUERNA

Coahuilaceratops or "Coahuila Horn Face," is a relatively new genus of Ornithischia Ceratopsidae, a  herbivorous ceratopsian dinosaur who lived during the Upper Cretaceous (late Campanian) near the town of Porvenir de Jalpa (about 64 km / 40 miles west of Saltillo) in what is now southern Coahuila (formerly Coahuila de Zaragoza), northern Mexico.

The Sierra Madre Oriental mountain range runs northwest to southwest forming a spine through the centre of the State. East of the range, the arid landscape slopes gently through the desert terrain down to the Rio Grande. It is home to wonderful common, rare and endangered cacti, beautiful (and one of my favourite) raptors, Aquila chrysaetos and the evolutionarily unlikely pronghorn, Antilocapra americana (if a monkey/owl/ antelope had a baby...)

The world was a much wetter warmer place when these big beauties roamed. Picture them ambling through lush vegetation and rearing young next to freshwater rivers, brackish swamps and salty ancient seas. Many of the dinosaur remains from the area bear the marks or remains of fossilized snails and clams. Perhaps predation vs a symbiotic relationship as proximity isn't always intimacy. Coahuilaceratops magnacuerna is known from holotype CPC 276, a partial skeleton of an adult along with bits and pieces of skull, a section of horn, pretty complete lower jaw, a smidge of the upper jaw and part of the frill.

Another specimen, CPS 277, has been touted as a possible juvenile Coahuilaceratops. All the specimens from Coahuilaceratops come from a single Upper Cretaceous (Campanian) locality of the Cerro del Pueblo Formation, northern Mexico.

This particular species of Coahuilaceratops was formally named C. magnacuerna by Mark A. Loewen, Scott D. Sampson, Eric K. Lund, Andrew A. Farke, Martha C. Aguillón-Martínez, C.A. de Leon, R.A. Rodríguez-de la Rosa, Michael A. Getty and David A. Eberth in 2010. Though the name was in circulation informally by those working in the study of ceratopsian dinosaurs as early as 2008.

Though challenged by examining and interpreting mere bits and pieces, the team posed estimates on the overall size of this new rather largish, 6.7 m / 22 ft, chasmosaurine. Coahuilaceratops' horns are also impressively large, estimated at 1.2 m / 4 feet. Rather long for a ceratopsian (consider that a Triceratops distinctive horn generally comes in under 115 cm / 45 inches and interesting in terms of evolutionary design. The holotypes are available for viewing at the Museo del Desierto in Saltillo, Coahuila. Photo credit: José F. Ventura

Thursday 4 January 2024

UNESCOCERATOPS KOPPELHUSAE BY JULIUS CSOTONYI

Unescoceratops koppelhusae, Julius Csotonyi
A very sweet small leptoceratopsid dinosaur, Unescoceratops koppelhusae — a new species in the collections of the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta.

The colourful and beautifully detailed painting you see here is by the very talented Julius Csotonyi who captured the magnificence of form, texture and palette to bring this small leptoceratopsid dinosaur to life.

The Royal Tyrrell Museum of Palaeontology, named in honour of Joseph Burr Tyrrell, is a palaeontology museum and research facility in Drumheller, Alberta, Canada. 

This jaw is the holotype specimen of this small leptoceratopsid dinosaur. Only a handful of isolated fossils have been found from this species, including a jaw that is the holotype specimen now in collections at the Royal Tyrell. 

The Royal Tyrrell Museum of Palaeontology, named in honour of Joseph Burr Tyrrell, is a palaeontology museum and research facility in Drumheller, Alberta, Canada. 

Unescoceratops koppelhusae, RTMP Collections
The rusty chocolate jaw bone you see here is the puzzle piece that helped all of the research come together and help us to better understand more about the diminutive leptoceratopsid dinosaurs from Alberta. 

The Cleveland Museum of Natural History's Michael Ryan and David Evans of the Royal Ontario Museum in Toronto recently determined that the specimen was a new genus and species. 

Unescoceratops is a genus of leptoceratopsid ceratopsian dinosaurs known from the Late Cretaceous (about 76.5-75 million years ago) of Alberta, Canada. Unescoceratops is thought to have been between one and two meters long and less than 91 kilograms. A plant-eater, its teeth were the roundest of all Leptocertopsids.

Dinosaur Provincial Park, Alberta, Canada
The genus name acknowledges the UNESCO  World Heritage Site, Dinosaur Provincial Park, where the fossil was found. 

In addition to its particularly beautiful scenery, Dinosaur Provincial Park – located at the heart of the province of Alberta's badlands – is unmatched in terms of the number and variety of high-quality specimens.

To date, they represent more than 44 species, 34 genera and 10 families of dinosaurs, dating back 75-77 million years. This provides us with remarkable insight into life millions of years ago.

The park contains exceptional riparian habitat features as well as badlands of outstanding aesthetic value.

The creamy honey, beige and rust coloured hills around the fossil locality are outstanding examples of major geological processes and fluvial erosion patterns in semi-arid steppes — think glorious! 

The scenic badlands stretch along 26 kilometres of high quality and virtually undisturbed riparian habitat, presenting a landscape of stark but exceptional natural beauty.

The species name honours Dr. Eva Koppelhus, who has made significant contributions to vertebrate palaeontology and palynology. 

The genus is named to honour the UNESCO World Heritage Site designation for the locality where the specimen was found and from the Greek “ceratops,” which means 'horned face'. 

Dr Michael Ryan explained that he meant to honour UNESCO's efforts to increase understanding of natural history sites around the world.

© Julius T. Csotonyi An illustration of Unescoceratops koppelhusae, a plant-eating dinosaur from the Late Cretaceous period that lived approximately 75 million years ago shared with his gracious permission. 

ABOUT THE ARTIST

Dr. Julius Csotonyi is a Vancouver-based scientific illustrator and natural history fine artist. He is a featured paleoartist on Season One and Season Two of BC's Fossil Bounty. Julius has a scientific background in ecology (MSc) and microbiology (PhD) which has taken him to study sensitive ecosystems, from sand dunes in the Rocky Mountain parks to hydrothermal vents at the bottom of the Pacific Ocean. 

These experiences have fuelled his strong resolve to work toward preserving our Earth’s biota. Painting biological subjects is one means that he uses to both enhance public awareness of biological diversity and to motivate concern for its welfare.   

He paints murals and panels that have appeared in numerous museums including the Smithsonian’s National Museum of Natural History, press release images for scientific publications, books, stamp sets — including the outstanding 2018 “Sharks of Canada” set for Canada Post — and coins for the Royal Canadian Mint. To view more of Julius Csotonyi's exquisite work visit: https://csotonyi.com/

Wednesday 27 December 2023

SVALBARD: ICE, SNOW AND ICHTHYOSAURS

Reindeer, Rangifer tarandus 
Ho Ho Ho. Ice, Snow, Reindeer & Ichthyosaurs — Svalbard is just what I imagine my version of Valhalla to be like, without all the mead, murder and mayhem. 

This Norwegian archipelago sits between mainland Norway and the North Pole. 

One of the world’s northernmost inhabited areas, it is known for its rugged, remote terrain of glaciers and frozen tundra sheltering polar bears, reindeer and Arctic fox. 

It is also known for reindeer. The lovelies you see here are all females as the males lose their antlers in the winter. So Rudolf and the rest of Santa's crew who pull his sleigh for him would have all been females as they are pictured with antlers. They are also shown flying across the sky, so the science gets a bit creative.

The Northern Lights or Nordlys are visible during winter, and summer brings the Midnight Sun — sunlight 24 hours a day. Norway or Norge is one of the very few locations where sunset merges into the sunrise, with no darkness in between, creating a soft, captivating twilight in which to view the world. 

The Botneheia Formation is made up of dark grey, laminated shales coarsening upwards to laminated siltstones and sandstones. South of the type area, the formation shows four coarsening-upward units. 

The formation is named for Botneheia Mountain, a mountain in Nordenskiöld Land at Spitsbergen, Svalbard. It has a height of 522 m.a.s.l., and is located south of Sassenfjorden, east of the valley of De Geerdalen. 

Svalbard, Norway
I was asked recently if folk head out in the torrential rain or ice and snow to fossil collect. I would generally say yes for those where the potential prize always outweighs the weather. For Svalbard, it is a resounding yes. 

You have to remove the snow cover — or ice if you are impatient or unlucky — to get to the outcrops here. It is well worth the effort. Beneath the icy cover, you find lovely ammonoids and bivalves. 

Tastier still, ichthyosaur remains are found here. The first Triassic ichthyosaurs from Svalbard were found in the early 20th century. Now there are quite a few Triassic and Jurassic ichthyosaur species from this archipelago.

Two ichthyosaur specimens have been recovered that are of particular interest. They comprise part of the trunk and the caudal vertebral column respectively. 

Some features, such as the very high and narrow caudal and posterior thoracic neural spines, the relatively elongate posterior thoracic vertebrae and the long and slender haemapophyses indicate that they probably represent a member of the family Toretocnemidae. 

Ichthyosaur Bones
Numerous ichthyosaur finds are known from the underlying Lower Triassic Vikinghøgda Formation and the overlying Middle to Upper Triassic Tschermakfjellet Formation, the new specimens help to close a huge gap in the fossil record of the Triassic ichthyosaurs from Svalbard. 

There is a resident research group working on the Triassic ichthyosaur fauna, the Spitsbergen Mesozoic Research Group. 

Lucky for them, they often find the fossil remains fully articulated — the bones having retained their spatial relationship to one another. 

Most of their finds are of the tail sections of primitive Triassic ichthyosaurs. In later ichthyosaurs, the tail vertebrae bend steeply downwards and have more of a fish-like look. 

In these primitive ancestors, the tail looks more eel-like — bending slightly so that the spines on the vertebrae form more of the tail. 

Maisch, Michael W. and Blomeier, Dierk published on these finds back in 2009: Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen Band 254 Heft 3 (2009), p. 379 - 384. Nov 1, 2009.

Svalbard, Norway (Norge)
Svalbard was so remote that there were no Inuit or First Nation settlements. It is certainly possible an earlier people came through these islands, but they did not leave any trace of their travels. 

The first documented travellers to explore Spitsbergen arrived in 1795 as part of a hunting expedition. They included people from the arctic town of Hammerfest in Norway's far north. They were an excellent choice as they were used to barren, inhospitable lands and sailed to discover more. 

We know them as the Coast Sámi — a hearty, rugged people probably best known in history for their chieftain, Ottar. He left Hammerfest in the 9th century to visit then join King Alfred the Great's court in a newly forming England. 

Expeditions to the remote islands of Svalbard continued into the early 1800s and finally, a settlement was eked out of the cold landscape and slowly expanded to the rest of the islands. While today the islands are called Svalbard, I would have named them for the Norwegian word for remote — fjernkontroll.

Aristoptychites euglyphus and Daonella sp.
This marvellous block is filled with Aristoptychites (syn = Arctoptychites) euglyphus (Mojsisovics, 1886) and Daonella sp., oyster-like clams or bivalves from the Middle Triassic, Ladinian, rugged windswept outcrops at the top of the Daonella Shales, Botneheia Formation, Spitzbergen, Edgeøya and Barentsøya, eastern Svalbard, Norway. 

Daonella and Monotis are important species for our understanding of biostratigraphy in the Triassic and are useful as Index fossils. 

Index fossils are fossils used to define and identify geologic periods or faunal stages. To be truly useful, they need to have a short vertical range, wide geographic distribution and rapid evolutionary development.

Daonellids preferred soft, soupy substrates and we tend to find them in massive shell beds. Generally, if you find one, you find a whole bunch cemented together in coquina. The lovely block you see here is in the collections of the deeply awesome John Fam. 

Learning Languages

The Sámi languages (/ˈsɑːmi/ SAH-mee), Sami or Saami, are a group of Uralic languages spoken by the Sámi people in Northern Europe in parts of northern Finland, Norway, Sweden, and extreme northwestern Russia. Of the world's languages, I find them the most difficult for my mind and tongue to wrap around. The Uralic languages will be familiar to you as Hungarian (Magyar nyelv), Finnish and Estonian. 

Since my Sámi is terrible, I will share a few words of Norwegian that may come in handy if you visit Svalbard and have a hankering for their tasty fossils or fossiler. To say, ice, snow, reindeer and ichthyosaurs in Norwegian, you would say: is, snø, reinsdyr og ikthyosaurer

To say, "hello, where can I find fossils?" Use, "Hei, hvor kan jeg finne fossiler?" An expression you may not need but circumstances being what they are, "That is a big polar bear," is "Det er en stor isbjørn." A solid follow-up would be, "nice bear, run..." as "Fin bjørn, løp..." Good luck with that.

Wishing you and yours the very best of the holidays however you celebrate. 

Thursday 8 June 2023

VANCOUVER ISLAND'S FABULOUS FOSSILS: TRENT RIVER PALAEONTOLOGY

Dan Bowen, Chair, VIPS, Trent River
The rocks that make up the Trent River on Vancouver Island are on the move. They were laid down near of the equator as small, tropical islands. They rode across the Pacific heading north and slightly east over the past 85 million years to where we find them today.

The Pacific Plate is an oceanic tectonic plate that lies beneath the Pacific Ocean. And it is massive. At 103 million km2 (40 million sq mi), it is the largest tectonic plate and continues to grow fed by volcanic eruptions that piggyback onto its trailing edge.

This relentless expansion pushes the Pacific Plate into the North American Plate. The pressure subducts it beneath our continent where it then melts back into the earth. Plate tectonics are slow but powerful forces. 

The island chains that rode the plates across the Pacific smashed into our coastline and slowly built the province of British Columbia. And because each of those islands had a different origin, they create pockets of interesting and diverse geology.

It is these islands that make up the Insular Belt — a physio-geological region on the northwestern North American coast. It consists of three major island groups — and many smaller islands — that stretches from southern British Columbia up into Alaska and the Yukon. These bits of islands on the move arrived from the Late Cretaceous through the Eocene — and continues to this day.

The rocks that form the Insular Superterrane are allochthonous, meaning they are not related to the rest of the North American continent. The rocks we walk over along the Trent River are distinct from those we find throughout the rest of Vancouver Island, Haida Gwaii, the rest of the province of British Columbia and completely foreign to those we find next door in Alberta.

To discover what we do find on the Trent takes only a wee stroll, a bit of digging and time to put all the pieces of the puzzle together. The first geological forays to Vancouver Island were to look for coal deposits, the profitable remains of ancient forests that could be burned to the power industry.

Jim Monger and Charlie Ross of the Geological Survey of Canada both worked to further our knowledge of the complex geology of the Comox Basin. They were at the cutting edge of west coast geology in the 1970s. It was their work that helped tease out how and where the rocks we see along the Trent today were formed and made their way north.

We know from their work that by 85 million years ago, the Insular Superterrane had made its way to what is now British Columbia. 

The lands were forested much as they are now but by extinct genera and families. The fossil remains of trees similar to oak, poplar, maple and ash can be found along the Trent and Vancouver Island. We also see the lovely remains of flowering plants such as Cupanities crenularis, figs and breadfruit.

Heading up the river, you come to a delineation zone that clearly marks the contact between the dark grey marine shales and mudstones of the Haslam Formation where they meet the sandstones of the Comox Formation. Fossilized material is less abundant in the Comox sandstones but still contains some interesting specimens. Here you begin to see fossilized wood and identifiable fossil plant material.

Further upstream, there is a small tributary, Idle Creek, where you can find more of this terrestrial material in the sandy shales. As you walk up, you see identifiable fossil plants beneath your feet and jungle-like, overgrown moss-covered, snarly trees all around you.

Walking west from the Trent River Falls at the bottom, you pass the infamous Ammonite Alley, where you can find Mesopuzosia sp. and Kitchinites sp. of the Upper Cretaceous (Santonian), Haslam Formation. Minding the slippery green algae covering some of the river rocks, you can see the first of the Polytychoceras vancouverense zone.

Continuing west, you reach the first of two fossil turtle sites on the river — amazingly, one terrestrial and one marine. If you continue, you come to the Inland Island Highway.

The Trent River has yielded some very interesting marine specimens, and significant terrestrial finds. We have found a wonderful terrestrial helochelydrid turtle, Naomichelys speciosa, and the caudal vertebrae of a Hadrosauroid dinosaur. Walking down from the Hadrosaur site you come to the site of the fossil ratfish find — one of the ocean's oddest fish.

Ratfish, Hydrolagus Collie, are chimaera found in the north-eastern Pacific Ocean today. The fossil specimen from the Trent would be considered large by modern standards as it is a bruiser in comparison to his modern counterparts. 

This robust fellow had exceptionally large eyes and sex organs that dangled enticingly between them. You mock, but there are many ratfish who would differ. While inherently sexy by ratfish standards, this fellow was not particularly tasty to their ancient marine brethren (or humans today) — so not hugely sought after as a food source or prey.

A little further again from the ratfish site we reach the contact of the two Formations. The rocks here have travelled a long way to their current location. With them, we peel away the layers of the geologic history of both the Comox Valley and the province of British Columbia.

The Trent River is not far from the Puntledge, a river whose banks have also revealed many wonderful fossil specimens. The Puntledge is also the name used by the K'ómoks First Nation to describe themselves. They have lived here since time immemorial. Along with Puntledge, they refer to themselves as Sahtloot, Sasitla and Ieeksun.

References: Note on the occurrence of the marine turtle Desmatochelys (Reptilia: Chelonioidea) from the Upper Cretaceous of Vancouver Island Elizabeth L. Nicholls Canadian Journal of Earth Sciences (1992) 29 (2): 377–380. https://doi.org/10.1139/e92-033; References: Chimaeras - The Neglected Chondrichthyans". Elasmo-research.org. Retrieved 2017-07-01.

Directions: If you're keen to explore the area, park on the side of Highway 19 about three kilometres south of Courtenay and hike up to the Trent River. Begin to look for parking about three kilometres south of the Cumberland Interchange. There is a trail that leads from the highway down beneath the bridge which will bring you to the Trent River's north side.

Sunday 16 October 2022

VANCOUVER ISLAND'S HASLAM FORMATION

Steller's Jay, Cyanocitta stelleri
One of the classic Vancouver Island fossil localities is the Santonian-Maastrichtian, Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake, Nanaimo, British Columbia, Canada.

The quarry is no longer active as such though there is a busy little gravel quarry a little way down the road closer to Ammonite falls near Benson Creek Falls.

Today it is an active motocross site and remains one of the classic localities of the Nanaimo Group. We find well-preserved nautiloids and ammonites — Canadoceras, Pseudoschloenbachia, Epigoniceras — the bivalves — Inoceramus, Sphenoceramus— gastropods, and classic Nanaimo Group decapods — Hoploparia, Linuparus. We also find fossil fruit and seeds which tell the story of the terrestrial history of Vancouver Island.

Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake
It was John Fam, Vice-Chair, Vancouver Island Paleontological Society (VanPS), who originally told me about the locality. John is one of the most delightful and knowledgeable people you'd be well-blessed to meet.

While he lived on Vancouver Island, he was an active member of the VanPS back when I was Chair. Several of the best joint VIPS/VanPS paleontological expeditions were planned with or instigated by his passion for fossils. I tip my hat to him for his passion and shared love of all things paleo.

John grew up 15 minutes from the motocross locality and used to collect there a few times a week with his father. John has wonderful parents and since marrying his childhood sweetheart, the amazing Grace, those excellent genetics, curiosity and love of fossils are now being passed to a new generation. It's lovely to see John and Grace continuing tradition with two boys of their own.

I met John way back then and did an overnight at his parent's house the Friday before a weekend field trip to Jurassic Point. It was a joy to have him walk me through his collections and tell his stories from earlier years. After learning about the site from John, I headed up to the Motocross Pit with my Uncle Doug. He was a delightful man who grew up on the coast and had explored much of it but not the fossil site just 10-minutes from his home. It was wonderful to walk through time with him so many years ago and then again solo this past year with sadness in my belly that one of the best I've ever known has left this Earth.

Upper Cretaceous Haslam Formation Motocross Pit near Brannen Lake
There were some no trespassing signs up but no people around, so I walked the periphery looking for the bedrock of the Haslam.

The rocks we find here were laid down south of the equator as small, tropical islands. They rode across the Pacific heading north and slightly east over the past 80 million years to where we find them today.

Jim Haggart and Peter Ward have done much to increase our understanding of the molluscan fauna of the Nanaimo Group. Personally, both personify the charming Indiana Jones school of rugged manly palaeontologists you picture in popular film. Professionally, their singular contributions and collaborative efforts have helped shape our understanding of the correlation of Nanaimo Group fauna to those we find in the Gulf Islands of British Columbia and down in the San Juan Islands of Washington State.

Their work builds on the work of Usher (1952), Matsumoto (1959a, 1959b) and Mallory (1977). A healthy nod goes out to the work of Muller and Jeletzky (1970) for untangling the lithostratigraphic and biostratigraphic foundation for our knowledge of the Nanaimo Group.

Candoceras yokoyama, Photo: John Fam, VanPS
As I walked along the bedrock of the Haslam, a Steller's Jay, Cyanocitta stelleri, followed me from tree to tree making his guttural shook, shook, shook call. Instructive, he seemed to be encouraging me, timing his hoots to the beat of my hammer. Vancouver Island truly has glorious flora and fauna.

Fancy some additional reading? Check out a paper published in the Journal of Paleontology back in 1989 by Haggard and Ward on new Nanaimo Group Ammonites from British Columbia and Washington State.

In it, they look at the ammonite species Puzosia (Mesopuzosia) densicostata Matsumoto, Kitchinites (Neopuzosia) japonicus Spath, Anapachydiscus cf. A. nelchinensis Jones, Menuites cf. M. menu (Forbes), Submortoniceras chicoense (Trask), and Baculites cf. B. boulei Collignon are described from Santonian--Campanian strata of western Canada and northwestern United States.

Stratigraphic occurrences and ranges of the species are summarized and those taxa important for correlation with other areas in the north Pacific region and Late Cretaceous ammonite fauna of the Indo-Pacific region. Here's the link: https://www.jstor.org/stable/1305358?seq=1

Peter Ward is a prolific author, both of scientific papers and more popularized works. I highly recommend his book Gorgon: Paleontology, Obsession, and the Greatest Catastrophe in Earth's History. It is an engaging romp through a decade's research in South Africa's Karoo Desert.

Photo: Candoceras yokoyamai from Upper Cretaceous Haslam formation (Lower Campanian) near Nanaimo, British Columbia. One of the earliest fossils collected by John Fam (1993). Prepared using only a cold chisel and hammer. Photo & collection of John Fam, VIPS.

Sunday 2 October 2022

TRUMPET CALLS FROM THE CRETACEOUS

Reconstruction of Prosaurolophus maximus
When this good looking fellow was originally described by Brown, Prosaurolophus maximus was known only from a skull and jaw. Half of the skull was badly weathered at the time of examination, and the level of the parietal was distorted and crushed upwards to the side. 

You can imagine that these deformations in preservation created some grief in the final description.

Prosaurolophus maximus was a large-headed duckbill dinosaur, or hadrosaurid, in the ornithischian family Hadrosauridae.

The most complete Prosaurolophus maximus specimen had a massive skull an impressive 0.9 metres (3.0 ft) long that graced a skeleton about 8.5 metres (28 ft) long. 

He had a small, stout, triangular crest in front of his eyes. The sides of the crest are concave, forming depressions. 

The crest grew isometrically — without changing in proportion — throughout the lifetime of each individual, leading one to wonder if Prosaurolophus had had a soft tissue display structure such as inflatable nasal sacs. We see this feature in hooded seals, Cystophora cristata, who live in the central and western North Atlantic today. Prosaurolophus maximus may have used their inflatable nasal sac for a display to warn a predator or to entice the ladies, attracting the attention of a female.

The different bones of the skull are easily defined with the exception of the parietal and nasal bones. Brown found that the skull of the already described genus Saurolophus was very similar overall, just smaller than the skull of Prosaurolophus maximus. The unique feature of a shortened frontal in lambeosaurines is also found in Prosaurolophus maximus, and the other horned hadrosaurines Brachylophosaurus, Maiasaura, and Saurolophus. Although they lack a shorter frontal, the genera Edmontosaurus and Shantungosaurus share an elongated dentary structure.

Prosaurolophus maximus, Ottawa Museum of Nature
Patches of preserved skin are known from two juvenile specimens, TMP 1998.50.1 and TMP 2016.37.1; these pertain to the ventral extremity of the ninth through fourteenth dorsal ribs, the caudal margin of the scapular blade, and the pelvic region. 

Small basement scales (scales that make up the majority of the skin surface), 3–7 millimetres (0.12–0.28 in) in diameter, are preserved on these patches - this is similar to the condition seen in other saurolophine hadrosaurs.

More uniquely, feature scales (larger, less numerous scales which are interspersed within the basement scales) around 5 millimetres (0.20 in) wide and 29 millimetres (1.1 in) long are found interspersed in the smaller scales in the patches from the ribs and scapula (they are absent from the pelvic patches). 

Similar scales are known from the tail of the related Saurolophus angustirostris (on which they have been speculated to indicate pattern), and it is considered likely adult Prosaurolophus would've retained the feature scales on their flanks like the juveniles.

Image: Three-dimensional reconstruction of Prosaurolophus maximus. Created using the skull reconstructions in the original description as reference. (Fig. 1 and 3 in Brown 1916). According to Lull and Wright (1942), the muzzle was restored too long in its original description. The colours and/or patterns, as with nearly all reconstructions of prehistoric creatures, are speculative. Created & uploaded to Wikipedia by Steveoc 86.

Wednesday 28 September 2022

TRACKING DINOSAURS

Stepping back in time can prove a challenge for the modern Time Traveller. But sometimes, if we're lucky, there are footsteps to follow.

This is a theropod track left by an Allosaurus walking through a muddy bank that turned to stone. These big beasties were large carnosaurian theropod dinosaurs that lived 155 to 145 million years ago.

This track and many others are at Dinosaur Ridge, a segment of the Dakota Hogback in the Morrison Fossil Area National Natural Landmark located in Jefferson County, Colorado, near the town of Morrison and just west of Denver.

The Dinosaur Ridge area is one of the world's most famous dinosaur fossil localities. Along with the dinosaur bones, you find plant fossils, crocodile bones and a variety of reptile tracks. 

In 1877, fossil excavation began at Dinosaur Ridge under the direction of palaeontologist Othniel Charles Marsh. Some of the best-known dinosaurs were found here, including Stegosaurus, Apatosaurus, Diplodocus, and Allosaurus

In 1973, the area was recognized for its uniqueness as well as its historical and scientific significance when it was designated the Morrison Fossil Area National Natural Landmark by the National Park Service. In 1989, the Friends of Dinosaur Ridge was formed to address increasing concerns regarding the preservation of the site and to offer educational programs on the area's resources.

Visit Dinosaur Ridge Morrison Fossil Area National Natural Landmark

Fancy a trip? You can visit these wonderful tracks, the Dinosaur Ridge Exhibit Hall and walk the trails through the tracks. They have put up helpful interpretive signs that explain the local geology, a volcanic ash bed, trace fossils, paleo-ecology, economic development of coal, oil and clay, and many other geologic and paleontological features.

There are two trails and a visitor centre at Dinosaur Ridge. The visitor centre features information on trails and a small gift shop. If you do not pack a lunch, you'll want to visit the Stegosaurus Snack Shack located outside the visitor centre offering coffee, cold drinks, burritos, pretzels and more. It is open from 10am-4pm Mon-Sat and 11am-4pm on Sun from June through August. 

Check updates on their website concerning Covid-19 restrictions. This is a vast site with easy trails so it makes for a great family trip when museums and other indoor facilities are closed.

Know Before You Go

Uncover Colorado: https://www.uncovercolorado.com/landmarks/morrison-golden-fossil-areas-dinosaur-ridge/

Monday 19 September 2022

DIPPY RETURNS TO THE NATURAL HISTORY MUSEUM UK

This sexy monkey with the long neck lit in soft purple and blue is Dippy the Dinosaur.

Dippy was the first Diplodocus to ever go on display when it was gifted to the Natural History Museum over 100 years ago. It quickly became a star, capturing hearts and imaginations. 

In 1905 a cast of a Diplodocus skeleton was donated to the Museum by the wealthy businessman Andrew Carnegie, based on the original specimen in the Carnegie Museum in the USA.

King Edward VII had requested a copy of the newly discovered dinosaur after seeing a picture of it in Carnegie's Scottish castle. From 1979 to early 2017 the cast - known affectionately as Dippy - was on display in the Museum's Hintze Hall.

In 1993, Dippy's tail was lifted from the ground after research revealed that Diplodocus tails would have been raised high to balance the neck.

Every two years or so, Museum experts used specialist equipment to clean the 292 bones that makeup Dippy. It takes two staff members two days to clean the cast and make sure it is maintained for future generations to enjoy. 

Dippy left the Museum in 2017 to complete a whirlwind tour of the UK. Throughout his journey, Dippy witnessed the changing state of nature and how the UK's biodiversity is in sharp decline. The famous cast is now back visiting the Museum until Christmas 2022.

Diplodocus had a long neck that it would have used to reach high and low vegetation and to drink water. There has been some debate over how such a long neck would have been held.

Scientists now think that ligaments running from the hip to the back of the neck would have allowed Diplodocus to hold its neck in a horizontal position without using muscles. The vertebrae (back bones) are split down the middle and this space could have held ligaments like these. Diplodocus may have had narrow, pointed bony spines lining its back.

Sunday 18 September 2022

DARWIN'S TOXODON

Toxodon is an extinct large grazing mammal. The first Toxodon fossils were discovered by Charles Darwin on his visit to South America as part of his voyage on the HMS Beagle. 

Darwin wondered at the fossil's strange appearance as it seemed to share features with both rodents and rhinos. 

“Toxodon is perhaps one of the strangest animals ever discovered,” wrote Darwin. He first encountered the creature in Uruguay on November 26th, 1834. 

“Having heard of some giant’s bones at a neighbouring farm-house…, I rode there accompanied by my host, and purchased for the value of eighteen pence the head of the Toxodon.”

The beast’s skeleton, once fully assembled, was a baffling mish-mash of traits. It was huge like a rhino, but it had the chiselling incisors of a rodent—its name means “arched tooth”—and the high-placed eyes and nostrils of a manatee or some other aquatic mammal. “How wonderfully are the different orders, at present time so well separated, blended together in different points of the structure of the toxodon!”

Although Toxodon is not related to rodents, in 2015, it was discovered to be distantly related to rhinoceros. 

Tuesday 30 August 2022

SPINOSAURUS AEGYPTIACUS

Spinosaurus the Spine Lizard of the Cretaceous
This beautiful big boy painted in yellow, green and blue is Spinosaurus aegyptiacus. 

Spinosaurus (meaning "spine lizard") is a genus of spinosaurid dinosaur that lived in what now is North Africa during the Cenomanian to upper Turonian in the Late Cretaceous— 99 to 93.5 million years ago. 

The genus was known first from Egyptian remains discovered in 1912 and described by German palaeontologist Ernst Stromer in 1915. 

The original remains were destroyed in World War II, but additional material came to light in the early 21st century.  It is unclear whether one or two species are represented in the fossils reported in the scientific literature. The best known species is S. aegyptiacus from Egypt, although a potential second species, S. maroccanus, has been recovered from Morocco. The contemporary spinosaurid genus Sigilmassasaurus has also been synonymized by some authors with S. aegyptiacus, though other researchers propose it to be a distinct taxon. 

In 2014, Ibrahim and his colleagues suggested that Spinosaurus aegyptiacus could reach over 15 metres (49 ft) in length. In 2022, however, Paul Sereno and his colleagues suggested that Spinosaurus aegyptiacus reached a maximum body length of 14 metres (46 ft) and a maximum body mass of 7.4 metric tons (8.2 short tons) by constructing an adult flesh model "with the axial column in neutral pose."

Spinosaurus is the longest known terrestrial carnivore; other large carnivores comparable to Spinosaurus include theropods such as Tyrannosaurus, Giganotosaurus and Carcharodontosaurus. The most recent study suggests that previous body size estimates are overestimated, and that S. aegyptiacus reached 14 metres (46 ft) in length and 7.4 metric tons (8.2 short tons) in body mass. 

The skull of Spinosaurus was long, low, and narrow, similar to that of a modern crocodilian, and bore straight conical teeth with no serrations. It would have had large, robust forelimbs bearing three-fingered hands, with an enlarged claw on the first digit. The distinctive neural spines of Spinosaurus, which were long extensions of the vertebrae (or backbones), grew to at least 1.65 meters (5.4 ft) long and were likely to have had skin connecting them, forming a sail-like structure, although some have suggested that the spines were covered in fat to form hump.

Sunday 28 August 2022

SPLIT SAIL SPINOSAURID

Back in the Cretaceous in what is now the heart of the Indochinese Peninsula, a group of mighty spinosaur with an unusual split sail were playing and hunting in the local waters. 

Picture an alligator whose legs grew especially long, add a big split sail on his back and arms that could grasp. That's kind of what these big therapod dinosaurs looked like.

The first fossil remains of Ichthyovenator were found in 2010 at Ban Kalum in the Grès Supérieurs Formation of the Savannakhet Basin in Savannakhet Province in the province of Laos. We have found other fossil remains at the site including fossil fish, iguanodontians and some compelling sauropod and ornithopod tracks.

Dinosaur remains are among the fossils that have been recovered from the formation. It is equivalent to the Khok Kruat Formation of northeastern Thailand, a fluvial formation similar in age. 

The Khok Kruat Formation is a treasure trove of fossil goodies as well. We have found the remains of fossil sharks, fish, turtles, crocodilians and pterosaurs. Phuwiangosaurus-like teeth, Siamosaurus teeth, and fragmentary postcranial remains of spinosaurids have been dug up there. We have also found Fukuiraptor-like teeth—and, being close in age, they are both Albian, it is likely just a matter of time before we find Ichthyovenator bones there as well.

Ichthyovenator had a long, shallow snout and robust forelimbs. Its diet likely mainly consisted of aquatic prey, hence its etymology. Spinosaurids are also known to have eaten small dinosaurs and pterosaurs in addition to fish. 

Ichthyovenator's conspicuous split sail might have been used for sexual display or species recognition. Fossil evidence suggests spinosaurids, especially spinosaurines, were adapted for semiaquatic lifestyles. The vertebral spines of Ichthyovenator's tail were unusually tall, suggesting—as in today's crocodilians—the tail may have aided in swimming. Ichthyovenator lived alongside sauropod and ornithopod dinosaurs, as well as bivalves, fish and turtles.

These fossilized bones were recovered from a red sandstone layer within a surface area of less than 2 square metres (22 sq ft). 

So, what did we find? So many great bits and pieces of these giant beasts as a massive jigsaw puzzle. Gathered together, they have been designated under the specimen numbers MDS BK10-01 to 15. There is a partly articulated, well-preserved skeleton. Sadly, missing are the skull and limbs. In with the bits and pieces were the third-to-last dorsal (back) vertebra, the neural spine of the last dorsal vertebra, five partial sacral (hip) vertebrae, the first two caudal (tail) vertebrae, both ilia (main hip bones), a right pubis (pubic bone), both ischia (lower and rearmost hip bones) and a posterior dorsal rib. 

The twelfth dorsal spine is bent sideways when viewed front-to-back due to taphonomic distortion. The central (vertebral bodies) of the sacrals are largely incomplete due to erosion but preserved all of their accompanying spines with their upper edges intact. 

At the time of Ichthyovenator's description, excavations at the site were still ongoing. Cast fossil spinal column laid out inside a glass display case at a museum Casts of the vertebrae at the Muséum National d'Histoire Naturelle, Paris After undergoing preparation in 2011, the skeleton was used as the basis, or holotype, for the type species Ichthyovenator laosensis, which was named and described in 2012 by palaeontologists Ronan Allain, Tiengkham Xeisanavong, Philippe Richir, and Bounsou Khentavong. 

The generic name is derived from the Old Greek word ἰχθύς (ichthys), "fish", and the Latin word venator, "hunter", in reference to its likely piscivorous or fish-eating lifestyle. The specific name refers to its provenance in Laos.

Ichthyovenator is the third named spinosaurid dinosaur from Asia after the Thai genus Siamosaurus in 1986 and the Chinese species "Sinopliosaurus" fusuiensis in 2009. The latter may represent the same animal as Siamosaurus

In 2014, Allain published a conference paper on Ichthyovenator; the abstract indicated additional remains from the original individual had been found after excavations continued in 2012. These remains include three teeth, the left pubis, and many vertebrae, including a nearly complete neck, the first dorsal vertebra, and seven more caudal vertebrae.

Some of these additional vertebrae were compared with those of other spinosaurids in a 2015 paper by German palaeontologist Serjoscha Evers and colleagues, in which they noted similarities with the vertebrae of the African spinosaurid Sigilmassasaurus

Allain and his team considered Ichthyovenator as representing the first unequivocal spinosaurid from Asia. Though prior spinosaurids had been named from the continent—including Siamosaurus from Thailand's Barremian Sao Khua Formation and "Sinopliosaurus" fusuiensis from China's Aptian Xinlong Formation—the authors noted that palaeontologists have debated the validity of these taxa because they are only confidently known from isolated teeth. 

Brazilian palaeontologists Marcos Sales and Caesar Schultz have suggested these teeth may eventually be attributed to spinosaurids similar to Ichthyovenator. In addition to tooth fossils, a spinosaurid skeleton that possibly belongs to Siamosaurus was excavated from the Thai Khok Kruat Formation in 2004 and was identified as a definite spinosaur in a 2008 conference abstract by Angela Milner and colleagues, eight years before Ichthyovenator's description.

Friday 19 August 2022

BOSS VERSUS HORN: PACHYRHINOSAURUS

The skulls of many dinosaurs had bumps, crests or horns for display, scaring rivals and impressing potential mates.

Instead of the sharp nose horn of many of their large plant-eating cousins, Pachyrhinosuaurs had a big, bony lump on their snout.

Pachyrhinosaurus were ceratopsian dinosaurs that lived in North America during the Late Cretaceous. 

Unlike their Triceratops cousins, Pachyrhinosaurus can be distinguished by their large lumps or bosses over their nose rather than horns.

The first fossil specimens were found in western Canada in Alberta by Charles M. Sternberg in 1946 and named in 1950. Charles Mortram Sternberg (1885–1981) was an American-Canadian fossil collector and palaeontologist, and son of the famous Charles Hazelius Sternberg (1850-1943) who worked with Edward Drinker Cope and Othniel C. Marsh, and for the British Museum, the San Diego Natural History Museum and other museums. 

Late in his career, Charles M. collected and described Pachyrhinosaurus, Brachylophosaurus, Parksosaurus and Edmontonia. A contemporary author wrote, "No published study of Canadian dinosaurs is possible today without citing one or another of Sternberg's papers. 

Over a dozen partial skulls and a large assortment of other fossils from various species have been found in Alberta and Alaska. A great number were not available for study until the 1980s, resulting in a relatively recent increase of interest in Pachyrhinosaurus.

Three species have been identified. P. lakustai, from the Wapiti Formation, the bonebed horizon of which is roughly equivalent age to the upper Bearpaw and lower Horseshoe Canyon Formations, is known to have existed from about 73.5–72.5 million years ago. P. canadensis is younger, known from the lower Horseshoe Canyon Formation, about 71.5–71 Ma ago and the St. Mary River Formation. 

Fossils of the youngest species, P. perotorum, have been recovered from the Prince Creek Formation of Alaska, and date to 70–69 Ma ago. The presence of three known species makes this genus the most speciose among the centrosaurines.

Thursday 18 August 2022

TOROSAURUS WALKING THROUGH THE FOG

Torosaurus was a ceratopsian dinosaur that lived during the Cretaceous. These fellows look very similar to their Triceratops cousins but are an entirely different species in their own right. 

In 1891, two years after the naming of Triceratops, a pair of ceratopsian skulls with elongated frills bearing holes were found in southeastern Wyoming, Niobrara County, by John Bell Hatcher. 

Hatcher's employer, palaeontologist Professor Othniel Charles Marsh, coined the genus Torosaurus for them. While an estimated 2,000 Triceratops specimens have been collected from the American West, only seven partial skulls of Torosaurus have been found, so they are pretty rare.

Like Triceratops, they had massive skulls. Torosaurus had one of the largest skulls of any known land animal, with the frilled skull reaching 2.77 metres (9.1 ft) in length. Torosaurus were about the same size as Triceratops. 

They had an elongated frill with large openings (fenestrae), long squamosal bones of the frill with a trough on their upper surface, and the presence of five or more pairs of hornlets (epoccipitals) on the back of the frill. Torosaurus also lacked the long nose horn seen in Triceratops prorsus, and instead resembled the earlier and more basal Triceratops horridus in having a short nose horn. Three species have been named, Torosaurus latus, T. gladius and T. utahensis. T. gladius is no longer considered a valid species.

Wyoming Outcrops
The individuals referred to as Torosaurus are all large, comparable to the largest Triceratops specimens. Due to the elongated frill, especially the skull length is considerable. Hatcher estimated the skull of YPM 1830 at 2.2 metres, and of YPM 1831 at 2.35 metres. 

In 1933, Richard Swann Lull increased this to 2.4 metres and 2.57 metres respectively. Based on this, Torosaurus was thought, albeit briefly, to have the longest skull of any known land animal. 

Sixty-five years later in 1998, Thomas Lehman claimed that a Pentaceratops specimen possessed a partial skull that would have been 2.9 metres long in life. This was again doubted by Nicholas Longrich who in 2011 named this exemplar as a separate genus Titanoceratops and concluded its skull had been reconstructed as too long.

In 2006, Andrew Farke, a palaeontologist at the Alf Museum of Paleontology in South Dakota, pointed out that the new skulls described by him were on average even longer than Hatcher's original two: MOR 1122 has a length of 252 centimetres and MOR 981 of 277 centimetres.

Farke’s research interests focus on exploring the Cretaceous continental ecosystems of North America, particularly the ceratopsian (horned) dinosaurs, with active fieldwork in California and Wyoming.

In 2006, Farke published some diagnostic traits of Torosaurus. The frill is extremely long in comparison to the remainder of the skull. The rear, parietal, and edge of the frill bear ten or more epiparietals, triangular osteoderms. A midline epiparietal is absent; likewise, no osteoderm straddles the parietal-squamosal boundary. The parietal bone is thin. It is pierced by parietal fenestrae in the form of circular or transversely oval openings. The parietal bone is about 20% wider than long. 

Farke also identified a single trait in which T. latus differed from both Triceratops horridus and T. utahensis: its squamosal bore a conspicuous ridge on the edge with the parietal combined with a deep longitudinal trough parallel to it.

Farke pointed out that the known Torosaurus specimens are rather variable. The orbital "brow" horns are sometimes large and curved to the front, as with MOR 981, and sometimes short and straight as shown by MOR 1122 and ANSP 15191. 

Also, the position of these horns differs: often they are located directly on top of the eye socket but with YPM 1831 they originate at the rear edge of the orbit. Likewise, there is a variation in the form of the nose horn. YPM 1831 and to a lesser extent YPM 1830 have a straight upright nasal horn but MOR 981, ANSP 15192 and especially MOR 1122 at most possess a low bump. The frill too differs. ANSP 15192 and YPM 1830 have a shield curving upwards at the rear, but the frill of YPM 1831 is nearly flat, though this could be an artefact of restoration. 

The frill of YPM 1831 is also heart-shaped, with a clear midline notch, whereas the rear edge of the other specimens is straight. The frill proportions are quite variable: with YPM 1831 the length-width ratio is 1.26 but MOR 981 has a shield 2.28 times longer than wide. The number of epiparietals is difficult to assess as most fossils seem to have lost them. MOR 981 and MOR 1122 have ten and twelve epiparietals respectively. YPM 1831 has been restored with a fontanelle in the skull roof, which possibly is authentic. Farke also concluded that the degree of variability did not exceed that shown by related genera.

Farke stressed that, apart from the frill, no systematic differences could be found between Torosaurus and Triceratops. All Torosaurus specimens are similar in that they lack a truly long nasal horn and a horizontal arterial groove at the front base of that horn, but Triceratops fossils with the same combination of traits are not uncommon. 

In 2008, Hunt concluded that T. utahensis, contrary to T. latus but similar to Triceratops, possessed a midline epiparietal.

Tuesday 16 August 2022

ARCHAEOPTERYX VS DRAGONFLY

An archaeopteryx, a Jurassic aged theropod dinosaur that looked like a skinny chicken, lunges at a dragonfly at the end of a smooth river stone.

Archaeopteryx were feathered dinosaurs and were once thought to be the oldest known fossil bird. 

We find their fossil remains in the Solnhofen Limestone outcrops that are more than 150 million years old.

The Solnhofen Plattenkalk, or Solnhofen Limestone, geologically known as the Altmühltal Formation, is a Jurassic Konservat-Lagerstätte that preserves a rare assemblage of fossilized organisms, including highly detailed imprints of soft bodied organisms such as sea jellies. 

The most familiar fossils of the Solnhofen Plattenkalk include the early feathered theropod dinosaur Archaeopteryx preserved in such detail that they are among the most famous and most beautiful fossils in the world. 

The Solnhofen beds lie in the German state of Bavaria (Bayern), halfway between Nuremberg (Nürnberg) and Munich (München) and were originally quarried as a source of lithographic limestone. The Jura Museum situated in Eichstätt, Germany has an extensive exhibit of Jurassic fossils from the quarries of Solnhofen and surroundings, including marine reptiles, pterosaurs, and one specimen of the early bird Archaeopteryx.

While all the first discoveries were from Bavaria, Germany, we have now found wonderfully preserved specimens in Liaoning, China, including Xiaotingia zhengi. The fossils we find in Germany are in a very fine-grained limestone that preserved their features with remarkable detail. 

These delightful proto-birds were preserved at the bottom of a shallow tropical marine lagoon whose lime-rich muds slowed their decay and encased their bones— and feathers— in mud. 

Sunday 14 August 2022

BIG HEAD LITTLE HOOVES: TRICERATOPS

This cutie is a Triceratops. The name means three-horned face in Greek but they might have been named Little Hooves instead as a nod to their weight-bearing fingers and toes that ended in sweet little hooves. 

Three of their five fingers and all of their toes end in a broad, flat-shaped hoof bone with a horny covering. 

Their hooves helped to protect their toes from wear and tear and support their heavy 5-ton bodies as they plodded along munching on cycads and palm fronds in the Late Cretaceous. These ceratopsid dinosaurs loved their plants. They used their beak-like jaws and slicing teeth to pluck and chew tasty foliage. Picture an animal about the size of an elephant, now lose the trunk, add the big head frill and horns. That's them!

Bearing a large bony frill, three horns on the skull, and a large four-legged body, exhibiting convergent evolution with rhinoceroses and bovines, Triceratops is one of the most recognizable of all dinosaurs and the most well-known ceratopsid. It was also one of the largest, up to 8–9 metres (26–30 ft) long and 5–9 metric tons (5.5–9.9 short tons) in body mass. 

It shared the landscape with and was most likely preyed upon by Tyrannosaurus, though it is less certain that two adults did battle in the fanciful manner often depicted in museum displays and popular images. The functions of the frills and three distinctive facial horns on its head have long inspired debate. Traditionally, these have been viewed as defensive weapons against predators. More recent interpretations find it probable that these features were primarily used in species identification, courtship, and dominance display, much like the antlers and horns of modern ungulates.

Triceratops was traditionally placed within the "short-frilled" ceratopsids, but modern cladistic studies show it to be a member of the Chasmosaurinae which usually have long frills. Two species, T. horridus and T. prorsus, are considered valid today, from the seventeen species that have ever been named. Research published in 2010 concluded that the contemporaneous Torosaurus, a ceratopsid long regarded as a separate genus, represents Triceratops in its mature form. This view has been disputed; further data is needed to settle the debate.

Triceratops has been documented by numerous remains collected since the genus was first described in 1889 by American paleontologist Othniel Charles Marsh. Specimens representing life stages from hatchling to adult have been found. As the archetypal ceratopsid, Triceratops is one of the most popular dinosaurs, and has been featured in film, postal stamps, and many other types of media.

Thursday 11 August 2022

TRUMPET CALLS FROM THE CRETACEOUS

Reconstruction of Prosaurolophus maximus
When this good looking fellow was originally described by Brown, Prosaurolophus maximus was known only from a skull and jaw. Half of the skull was badly weathered at the time of examination, and the level of the parietal was distorted and crushed upwards to the side. 

You can imagine that these deformations in preservation created some grief in the final description.

Prosaurolophus maximus was a large-headed duckbill dinosaur, or hadrosaurid, in the ornithischian family Hadrosauridae.

The most complete Prosaurolophus maximus specimen had a massive skull an impressive 0.9 metres (3.0 ft) long that graced a skeleton about 8.5 metres (28 ft) long. 

He had a small, stout, triangular crest in front of his eyes. The sides of the crest are concave, forming depressions. 

The crest grew isometrically — without changing in proportion — throughout the lifetime of each individual, leading one to wonder if Prosaurolophus had had a soft tissue display structure such as inflatable nasal sacs. We see this feature in hooded seals, Cystophora cristata, who live in the central and western North Atlantic today. Prosaurolophus maximus may have used their inflatable nasal sac for a display to warn a predator or to entice the ladies, attracting the attention of a female.

The different bones of the skull are easily defined with the exception of the parietal and nasal bones. Brown found that the skull of the already described genus Saurolophus was very similar overall, just smaller than the skull of Prosaurolophus maximus. The unique feature of a shortened frontal in lambeosaurines is also found in Prosaurolophus maximus, and the other horned hadrosaurines Brachylophosaurus, Maiasaura, and Saurolophus. Although they lack a shorter frontal, the genera Edmontosaurus and Shantungosaurus share an elongated dentary structure.

Prosaurolophus maximus, Ottawa Museum of Nature
Patches of preserved skin are known from two juvenile specimens, TMP 1998.50.1 and TMP 2016.37.1; these pertain to the ventral extremity of the ninth through fourteenth dorsal ribs, the caudal margin of the scapular blade, and the pelvic region. 

Small basement scales (scales that make up the majority of the skin surface), 3–7 millimetres (0.12–0.28 in) in diameter, are preserved on these patches - this is similar to the condition seen in other saurolophine hadrosaurs.

More uniquely, feature scales (larger, less numerous scales which are interspersed within the basement scales) around 5 millimetres (0.20 in) wide and 29 millimetres (1.1 in) long are found interspersed in the smaller scales in the patches from the ribs and scapula (they are absent from the pelvic patches). 

Similar scales are known from the tail of the related Saurolophus angustirostris (on which they have been speculated to indicate pattern), and it is considered likely adult Prosaurolophus would've retained the feature scales on their flanks like the juveniles.

Image: Three-dimensional reconstruction of Prosaurolophus maximus. Created using the skull reconstructions in the original description as reference. (Fig. 1 and 3 in Brown 1916). According to Lull and Wright (1942), the muzzle was restored too long in its original description. The colours and/or patterns, as with nearly all reconstructions of prehistoric creatures, are speculative. Created & uploaded to Wikipedia by Steveoc 86.